{"id":3687,"date":"2025-02-27T21:52:39","date_gmt":"2025-02-27T21:52:39","guid":{"rendered":"https:\/\/dornsife.usc.edu\/wrigley\/?p=3687"},"modified":"2026-04-02T22:51:53","modified_gmt":"2026-04-02T22:51:53","slug":"biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty","status":"publish","type":"post","link":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/","title":{"rendered":"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies"},"content":{"rendered":"\n\n\n\n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--article-hero \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--article-hero\"\n    \n      >\n\n    \n<div class=\"inner-wrapper\">\n          \n<div class=\"f--field f--image\">\n\n    \n    \n    \n    \n    \n    \n              \n      <img\n                            data-src=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-768x432.jpeg\"\n          data-srcset=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-1920x1080.jpeg 1920w,https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-1280x720.jpeg 1280w,https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-768x432.jpeg 768w\"          data-sizes=\"(min-width:1200px) 75vw, (min-width:768px) 83vw, 100vw\"          class=\"lazyload\"\n        \n                  alt=\"Karen Lloyd, wearing a dark blue beanie that says \"USC Wrigley Institute,\" smiles as she stands in a craggy, rocky landscape\"\n        \n        \n                                      \/>\n\n    \n    \n  \n  \n\n<\/div>\n  \n      <div class=\"image-caption\">\n          \n<div class=\"f--field f--description\">\n\n    \n  Biogeochemist Karen Lloyd on a research trip to Chile, where she traveled to study microbes that live in hot springs. (Photo: Karen Lloyd)\n\n\n<\/div>\n    <\/div>\n  \n  <div class=\"text-wrapper\">\n    \n              \n<div class=\"f--field f--page-title\">\n\n    \n  <h1>Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies<\/h1>\n\n\n<\/div>\n    \n    \n    \n          <span class=\"post-date-field\">February 27, 2025<\/span>\n      <\/div>\n<\/div>\n\n\n  <\/div><\/div>\n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--social-share \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--social-share\"\n    \n      >\n\n    \n  <div class=\"content-wrapper\">\n    <span class=\"a2a_kit a2a_kit_size_32 addtoany_list\" style=\"line-height: 32px;\">\n      <span class=\"title\">\n        Share\n      <\/span>\n                        <a class=\"a2a_button_copy_link\" target=\"_blank\" href=\"\/#copy_link\" rel=\"nofollow noopener\" title=\"Link\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_copy_link\">\n              <svg height=\"19\" viewBox=\"0 0 19 19\" width=\"19\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\"><path d=\"m7.43475275 9.52380952-2.17490843 2.26076008c-1.08745421 1.058837-1.68841575 2.518315-1.68841575 4.0350275 0 1.5167124.60096154 2.9475732 1.68841575 4.0350274 1.058837 1.0874543 2.51831502 1.6884158 4.03502747 1.6884158 1.44087681 0 2.80427251-.5423678 3.86936601-1.5289606l.1656615-.1594552 2.2321428-2.2321428-1.6311813-1.6311813-2.2321429 2.26076c-.6581959.658196-1.5167124 1.0016026-2.43246332 1.0016026s-1.7742674-.3434066-2.43246337-1.0016026c-.65819597-.6581959-1.00160257-1.5167124-1.00160257-2.4324633 0-.8547009.2991453-1.6595442.87416905-2.2981957l.12743352-.1342677 2.23214286-2.2321429zm6.88661855-.40736607-5.26116068 5.26116075 1.61881868 1.6188186 5.2611607-5.2611607zm1.3837569-5.54501488c-1.4408768 0-2.8042725.54236779-3.869366 1.52896062l-.1656615.15945513-2.23214282 2.23214286 1.63118132 1.63118132 2.2321429-2.23214286c.6581959-.65819597 1.5167124-1.00160256 2.4324633-1.00160256s1.7742674.34340659 2.4324634 1.00160256 1.0016026 1.51671246 1.0016026 2.43246337c0 .85470089-.2991453 1.65954419-.8741691 2.29819559l-.1274335.1342678-2.2321429 2.2321428 1.5739469 1.5739469 2.2321429-2.2321428c1.0874542-1.058837 1.6884157-2.5183151 1.6884157-4.03502751 0-1.51671245-.6009615-2.94757326-1.6884157-4.03502747-1.058837-1.08745421-2.518315-1.68841575-4.0350275-1.68841575z\" transform=\"translate(-3 -3)\"\/><\/svg>\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">Copy Link<\/span>\n          <\/a>\n                                <a class=\"a2a_button_facebook\" target=\"_blank\" href=\"\/#facebook\" rel=\"nofollow noopener\" title=\"Facebook\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_facebook\">\n              <svg height=\"19\" viewBox=\"0 0 19 19\" width=\"19\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\"><path d=\"m23.9512 6h-16.9024c-.57923625 0-1.0488.46956375-1.0488 1.0488v16.9024c0 .5792362.46956375 1.0488 1.0488 1.0488h9.10024v-7.34768h-2.46772v-2.87584h2.46772v-2.11508c0-2.45404 1.49872-3.79012 3.68828-3.79012.7389668-.00263323 1.4775241.03516995 2.21236.11324v2.56348h-1.51392c-1.18788 0-1.4174.56544-1.4174 1.39612v1.8316h2.84772l-.37088 2.87584h-2.47684v7.34844h4.83284c.5792362 0 1.0488-.4695638 1.0488-1.0488v-16.9024c0-.57923625-.4695638-1.0488-1.0488-1.0488z\" fill-rule=\"evenodd\" transform=\"translate(-6 -6)\"\/><\/svg>\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">Facebook<\/span>\n          <\/a>\n                                <a class=\"a2a_button_twitter\" target=\"_blank\" href=\"\/#twitter\" rel=\"nofollow noopener\" title=\"X\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_twitter\">\n              <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\"\n\t y=\"0px\" viewBox=\"0 0 30 30\" enable-background=\"new 0 0 30 30\" xml:space=\"preserve\" width=\"19\" height=\"19\">\n<path fill=\"#000\" fill-rule=\"evenodd\" clip-rule=\"evenodd\" d=\"M14.5,16.2l-0.8-1.1l-6.3-9h2.7l5.1,7.3l0.8,1.1\n\tl6.6,9.4h-2.7L14.5,16.2L14.5,16.2z M16.9,13.4l7.4-8.7h-1.8l-6.5,7.5L11,4.8H5l7.8,11.4L5,25.2h1.8l6.8-7.9l5.5,7.9h6L16.9,13.4\n\tL16.9,13.4z\"\/>\n<\/svg>\n\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">X<\/span>\n          <\/a>\n                                <a class=\"a2a_button_linkedin\" target=\"_blank\" href=\"\/#linkedin\" rel=\"nofollow noopener\" title=\"Linkedin\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_linkedin\">\n              <svg height=\"19\" viewBox=\"0 0 19 19\" width=\"19\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\"><path d=\"m25 21.4220779c0 1.974026-1.6038961 3.5779221-3.5779221 3.5779221h-11.84415582c-1.97402598 0-3.57792208-1.6038961-3.57792208-3.5779221v-11.84415582c0-1.97402598 1.6038961-3.57792208 3.57792208-3.57792208h11.84415582c1.974026 0 3.5779221 1.6038961 3.5779221 3.57792208zm-14.5584416-12.21428569c-.98701295 0-1.60389606.61688311-1.60389606 1.48051949-.12337663.8636364.49350649 1.4805195 1.48051946 1.4805195.987013 0 1.6038961-.6168831 1.6038961-1.4805195 0-.86363638-.6168831-1.48051949-1.4805195-1.48051949zm1.3571429 12.70779219v-8.6363636h-2.83766234v8.6363636zm10.2402597 0v-4.9350649c0-2.5909091-1.3571428-3.8246753-3.3311688-3.8246753-1.4805195 0-2.0974026.8636363-2.4675325 1.4805194v-1.2337662h-2.8376623v8.6363636h2.8376623v-4.9350649c0-.2467532 0-.4935065.1233767-.7402597.1233766-.4935065.6168831-.987013 1.4805194-.987013.987013 0 1.4805195.7402597 1.4805195 1.974026v4.564935z\" transform=\"translate(-6 -6)\"\/><\/svg>\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">LinkedIn<\/span>\n          <\/a>\n                                <a class=\"a2a_button_bluesky\" target=\"_blank\" href=\"\/#bluesky\" rel=\"nofollow noopener\" title=\"Bluesky\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_bluesky\">\n              <svg xmlns=\"http:\/\/www.w3.org\/2000\/svg\" width=\"50px\" height=\"50px\" viewBox=\"0 0 24 24\">\n                <path fill=\"#00000\" d=\"M12 10.8c-1.087-2.114-4.046-6.053-6.798-7.995C2.566.944 1.561 1.266.902 1.565C.139 1.908 0 3.08 0 3.768c0 .69.378 5.65.624 6.479c.815 2.736 3.713 3.66 6.383 3.364c.136-.02.275-.039.415-.056c-.138.022-.276.04-.415.056c-3.912.58-7.387 2.005-2.83 7.078c5.013 5.19 6.87-1.113 7.823-4.308c.953 3.195 2.05 9.271 7.733 4.308c4.267-4.308 1.172-6.498-2.74-7.078a8.741 8.741 0 0 1-.415-.056c.14.017.279.036.415.056c2.67.297 5.568-.628 6.383-3.364c.246-.828.624-5.79.624-6.478c0-.69-.139-1.861-.902-2.206c-.659-.298-1.664-.62-4.3 1.24C16.046 4.748 13.087 8.687 12 10.8\"  fill-rule=\"evenodd\" \/>\n              <\/svg>\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">Bluesky<\/span>\n          <\/a>\n                                <a class=\"a2a_button_email\" target=\"_blank\" href=\"\/#email\" rel=\"nofollow noopener\" title=\"E-mail\">\n            <span class=\"a2a_svg a2a_s__default a2a_s_print\">\n              <svg height=\"12\" viewBox=\"0 0 18 12\" width=\"18\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\"><path d=\"m8.14285714 9.42857143c-.17142857 0-.34285714 0-.51428571.08571428l7.28571427 6.34285719c.3428572.2571428.6857143.2571428.9428572 0l7.2857142-6.34285719c-.0857142-.08571428-.2571428-.08571428-.4285714-.08571428zm-1.28571428 1.11428567v.1714286 8.5714286c0 .6857143.6 1.2857143 1.28571428 1.2857143h14.57142856c.6857143 0 1.2857143-.6 1.2857143-1.2857143v-8.5714286c0-.0857143 0-.0857143 0-.1714286l-7.2 6.3428572c-.7714286.6857143-1.8857143.6857143-2.6571429 0z\" fill-rule=\"evenodd\" transform=\"translate(-6 -9)\"\/><\/svg>\n            <\/span>\n            <span class=\"a2a_label visually-hidden\">Email<\/span>\n          <\/a>\n                  <\/span>\n  <\/div>\n\n  <\/div><\/div>\n \n\n\n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n      \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><em>By Kathryn Royster<\/em><\/p>\n<p><span style=\"font-weight: 400;\"><a href=\"https:\/\/dornsife.usc.edu\/earth\/2024\/08\/08\/welcome-karen-lloyd\/\" target=\"_blank\" rel=\"noopener\">Karen Lloyd<\/a>, a renowned biogeochemist known for her work on microbes that live deep beneath the Earth\u2019s surface, has joined the USC faculty as the Wrigley Chair in Environmental Studies and professor of Earth sciences.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">&#8220;I am thrilled to welcome Karen to USC as part of the Wrigley Institute community and the university\u2019s world-class team of environmental and sustainability scholars,&#8221; says Wrigley Institute Director Joe \u00c1rvai. &#8220;She is the rare researcher who does groundbreaking work that addresses real-world needs while also communicating about it in a way that is both entertaining and insightful. The Wrigley Institute, and our extended community, are lucky to have her.\u201d<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Lloyd\u2019s research focuses on microbes that live in environments such as the bottom of the ocean, <a href=\"https:\/\/www.ted.com\/talks\/karen_lloyd_the_mysterious_microbes_living_deep_inside_the_earth_and_how_they_could_help_humanity\" target=\"_blank\" rel=\"noopener\">hot springs<\/a>, or under Arctic permafrost. Some of these microbes have extremely slow metabolisms\u2013so slow that <a href=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Lloyd-2020-time.pdf\" target=\"_blank\" rel=\"noopener\">they can go thousands of years<\/a> without reproducing, experiencing molecular turnover, or eating. Lloyd seeks to understand why the organisms are in this state, whether their slow metabolisms are an evolutionary advantage, what kinds of conditions might cause them to \u201cwake up,\u201d and how their presence and activities might intersect with climate change.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">To uncover the answers to these questions, Lloyd focuses heavily on a field-first approach to research. This approach comes partly by necessity: the microbes she studies reproduce so slowly that she could literally spend the rest of her life waiting for a culture to grow in a Petri dish. But she also prefers fieldwork for its true-to-life conditions.<\/span><\/p>\n<figure id=\"attachment_3690\" aria-describedby=\"caption-attachment-3690\" style=\"width: 800px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" class=\"wp-image-3690\" src=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond.jpg\" alt=\"two researchers working in a shallow, clear pond look small against a vast landscape of gray rock and brown hills\" width=\"800\" height=\"533\" srcset=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond.jpg 2048w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-300x200.jpg 300w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-1024x683.jpg 1024w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-768x512.jpg 768w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-1536x1024.jpg 1536w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-900x600.jpg 900w, https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Karen-Lloyd-Arctic-pond-1200x800.jpg 1200w\" sizes=\"(max-width: 800px) 100vw, 800px\" \/><figcaption id=\"caption-attachment-3690\" class=\"wp-caption-text\">Lloyd and another researcher collecting data at a hot spring in Chile. Lloyd studies slow-growing microbes that live for thousands, possibly even millions, of years in extreme environments such as hot springs, the Arctic permafrost, and the deep ocean. She prefers to study these microbes in the field as much as possible, to get an accurate understanding of how their physical environment and climate change affect them. (Photo: Jacopo Pasotti)<\/figcaption><\/figure>\n<p><span style=\"font-weight: 400;\">\u201cYou can do all the theoretical lab work you want, but when you\u2019re in the wild with so many factors affecting the organism, it may perform differently. That\u2019s why I always measure things in the wild as much as possible,\u201d Lloyd says.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Her preference for getting out into the field runs deep. Growing up on the rural North Carolina coast, Lloyd spent many hours exploring the environment around her home. She was particularly fond of the ocean\u2013so fond of it, in fact, that at age 13, she literally knocked on the door of the Duke University Marine Lab and asked if she could assist the researchers.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">\u201cI went to college with the aim of majoring in biology and just got sucked up by chemistry,\u201d Lloyd says. \u201cIt was like solving puzzles. But I couldn\u2019t imagine spending the rest of my life only in a lab, so I decided to study oceanography as well.\u201d<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Now that Lloyd has arrived in Los Angeles, the destination seems almost inevitable. She first encountered the Wrigley Institute and USC as a Ph.D. student, when she attended a summer course taught by faculty affiliate Will Berelson at the Wrigley Marine Science Center. That experience touched off a long-term connection with USC, even as she went on to a postdoctoral fellowship in Denmark and joined the faculty at the University of Tennessee Knoxville.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Over the years, she collaborated numerous times with the late <a href=\"https:\/\/dornsife.usc.edu\/news\/stories\/jan-amend-obituary\/\" target=\"_blank\" rel=\"noopener\">Jan Amend<\/a>, who was a microbiologist and USC Dornsife divisional dean of life sciences. Many of her other co-researchers have been USC graduate students or alumni. She says that USC\u2019s unique strengths in marine microbiology, geobiology, Earth sciences, and engineering were a strong draw for her. Add the wealth of local opportunities for collaboration\u2013with JPL, Caltech, UCLA, Cal State, and the Natural History Museum of L.A., to name a few\u2013and it was an easy \u201cyes\u201d to make the cross-country move from Knoxville to L.A.<\/span><\/p>\n<p>&nbsp;<\/p>\n<h3>VIDEO: How Karen Lloyd Finds Life in the Earth&#8217;s Deepest, Darkest Places<\/h3>\n<div class=\"video-embed\"><iframe loading=\"lazy\" width=\"560\" height=\"315\" src=\"https:\/\/www.youtube.com\/embed\/CBi9k4tkqxs?si=TxOBuQ7l-sLc3pk4\" title=\"Scientists Discover Thriving Life in Earth's Darkest Places\" frameborder=\"0\" allow=\"accelerometer; autoplay; clipboard-write; encrypted-media; gyroscope; picture-in-picture; web-share\" referrerpolicy=\"strict-origin-when-cross-origin\" allowfullscreen><\/iframe><\/div>\n<p><span style=\"font-weight: 400;\">Lloyd is excited to be in a coastal city once again, and to have the world\u2019s largest ocean (or, from her perspective, living lab) right on her doorstep. One of her first projects in her new role: studying methane-eating microbes in the San Pedro Channel between L.A. and Catalina Island.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Methane is a planet-warming gas that is up to 20 times more potent than carbon dioxide. It occurs naturally as plants and animals decay, but it\u2019s also created by human activity, especially the use of agricultural fertilizers. Methane in the San Pedro Channel comes from human and natural sources and is both produced in the ocean and carried to it by runoff.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">The gas is not naturally very soluble in water, so it tends to escape quickly into the atmosphere\u2013unless methane-eating microbes get to it first. Through their natural movements, or when they die, these microbes sequester the methane by taking it to deeper waters. Deep ocean water cycles to the surface slowly, so methane sequestered this way may stay out of the atmosphere for tens, hundreds, or even thousands of years.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">We don\u2019t currently know how much methane is in the San Pedro Channel, how much of it is escaping to the surface, or whether the local microbes are keeping up with the supply. Those are the questions Lloyd hopes to answer, aided by two Wrigley Institute postdoctoral researchers and the institute\u2019s <a href=\"https:\/\/dornsife.usc.edu\/spot\/\" target=\"_blank\" rel=\"noopener\">San Pedro Ocean Time-series<\/a>.\u00a0<\/span><\/p>\n<p><span style=\"font-weight: 400;\">Lloyd\u2019s results can help us understand the effects of runoff from large urban areas like Los Angeles, how quickly climate change may intensify as methane enters the atmosphere, and what we can do to protect ourselves and our planet. She also hopes her work will help people appreciate just how important these hidden, mysterious microbes are to our lives.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">\u201cThis kind of activity is happening not just in the channel but in groundwater, in the Arctic, in hot springs, and other places,\u201d Lloyd says. \u201cA huge biosphere lies under our feet all over the world, and we\u2019re just beginning to figure out what they\u2019re doing and how they can help us\u201d<\/span><\/p>\n<p><em>Karen Lloyd&#8217;s research on methane in the San Pedro Channel is supported in part by the Wrigley Institute&#8217;s Ballmer Group-funded Climate and Carbon Management Initiative.<\/em><\/p>\n<p><a href=\"https:\/\/dornsife.usc.edu\/wrigley\/research\/crosscutting-research-initiatives\/\">Learn more about the Climate and Carbon Management Initiative &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/dornsife.usc.edu\/wrigley\/research\/current-research-projects\/\">Learn more about Wrigley Institute faculty research &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/Lloyd-2020-time.pdf\" target=\"_blank\" rel=\"noopener\">Read Lloyd&#8217;s paper on slow-growing microbes &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/www.ted.com\/talks\/karen_lloyd_this_deep_sea_mystery_is_changing_our_understanding_of_life\" target=\"_blank\" rel=\"noopener\">Watch Lloyd&#8217;s TED Talk on deep-ocean microbes &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/gizmodo.com\/intraterrestrials-the-most-alien-life-weve-ever-seen-is-right-beneath-our-feet-2000599835\" target=\"_blank\" rel=\"noopener\">Read the Gizmodo review of Lloyd&#8217;s\u00a02025 book <em>Intraterrestrials<\/em> &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/blog.wfsu.org\/blog-coastal-health\/2025\/05\/deep-sea-food-webslife-on-the-ocean-floor-and-why-its-a-hard-place-to-explore-coast-to-canopy-episode-3\/\" target=\"_blank\" rel=\"noopener\">Listen to Lloyd&#8217;s appearance on WFSU Public Media, or read the transcript &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/www.noemamag.com\/theres-life-inside-earths-crust\/\" target=\"_blank\" rel=\"noopener\">Read Lloyd&#8217;s story about deep-Earth microbes on Noema &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/player.fm\/series\/sean-carrolls-mindscape-science-society-philosophy-culture-arts-and-ideas-2381982\/ep-314-karen-lloyd-on-the-deep-underground-biosphere\" target=\"_blank\" rel=\"noopener\">Listen to Lloyd&#8217;s appearance on the podcast <em>Sean Carroll&#8217;s Mindscape<\/em> &gt;&gt;<\/a><\/p>\n<p><a href=\"https:\/\/dornsife.usc.edu\/magazine\/buried-alive\/\" target=\"_blank\" rel=\"noopener\">Read &#8220;Buried Alive: The Secret Life of Deep Earth Microbes&#8221; (Dornsife Magazine) &gt;&gt;<\/a><\/p>\n\n\n\n<\/div>\n\n\n  <\/div><\/div>\n","protected":false},"excerpt":{"rendered":"","protected":false},"author":182,"featured_media":3688,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_acf_changed":false,"footnotes":""},"categories":[41],"tags":[45,72,16,71,73],"class_list":["post-3687","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-research","tag-climate-and-carbon-management-initiative","tag-earth-sciences","tag-faculty","tag-microbiology","tag-oceanography"],"acf":[],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.1.1 - https:\/\/yoast.com\/product\/yoast-seo-wordpress\/ -->\n<title>Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies<\/title>\n<meta name=\"description\" content=\"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.\" \/>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Karen Lloyd, who studies deep ocean microbes, joins USC faculty\" \/>\n<meta property=\"og:description\" content=\"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.\" \/>\n<meta property=\"og:url\" content=\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\" \/>\n<meta property=\"og:site_name\" content=\"Wrigley Institute for Environment and Sustainability\" \/>\n<meta property=\"article:published_time\" content=\"2025-02-27T21:52:39+00:00\" \/>\n<meta property=\"article:modified_time\" content=\"2026-04-02T22:51:53+00:00\" \/>\n<meta property=\"og:image\" content=\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg\" \/>\n\t<meta property=\"og:image:width\" content=\"2560\" \/>\n\t<meta property=\"og:image:height\" content=\"1920\" \/>\n\t<meta property=\"og:image:type\" content=\"image\/jpeg\" \/>\n<meta name=\"author\" content=\"Kathryn Royster\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\/\/schema.org\",\"@graph\":[{\"@type\":\"Article\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#article\",\"isPartOf\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\"},\"author\":{\"name\":\"Kathryn Royster\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e\"},\"headline\":\"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies\",\"datePublished\":\"2025-02-27T21:52:39+00:00\",\"dateModified\":\"2026-04-02T22:51:53+00:00\",\"mainEntityOfPage\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\"},\"wordCount\":12,\"commentCount\":0,\"image\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg\",\"keywords\":[\"climate and carbon management initiative\",\"Earth sciences\",\"faculty\",\"microbiology\",\"oceanography\"],\"articleSection\":[\"Research\"],\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"CommentAction\",\"name\":\"Comment\",\"target\":[\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#respond\"]}]},{\"@type\":\"WebPage\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\",\"url\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\",\"name\":\"Karen Lloyd, who studies deep ocean microbes, joins USC faculty\",\"isPartOf\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#website\"},\"primaryImageOfPage\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage\"},\"image\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg\",\"datePublished\":\"2025-02-27T21:52:39+00:00\",\"dateModified\":\"2026-04-02T22:51:53+00:00\",\"author\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e\"},\"description\":\"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.\",\"breadcrumb\":{\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#breadcrumb\"},\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"ReadAction\",\"target\":[\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/\"]}]},{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage\",\"url\":\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg\",\"contentUrl\":\"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg\",\"width\":2560,\"height\":1920,\"caption\":\"Karen Lloyd, wearing a dark blue beanie that says \\\"USC Wrigley Institute,\\\" smiles as she stands in a craggy, rocky landscape\"},{\"@type\":\"BreadcrumbList\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#breadcrumb\",\"itemListElement\":[{\"@type\":\"ListItem\",\"position\":1,\"name\":\"Home\",\"item\":\"https:\/\/dornsife.usc.edu\/wrigley\/\"},{\"@type\":\"ListItem\",\"position\":2,\"name\":\"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies\"}]},{\"@type\":\"WebSite\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#website\",\"url\":\"https:\/\/dornsife.usc.edu\/wrigley\/\",\"name\":\"Wrigley Institute for Environment and Sustainability\",\"description\":\"Creating a more sustainable and environmentally just future for our planet and all who live on it\",\"potentialAction\":[{\"@type\":\"SearchAction\",\"target\":{\"@type\":\"EntryPoint\",\"urlTemplate\":\"https:\/\/dornsife.usc.edu\/wrigley\/?s={search_term_string}\"},\"query-input\":{\"@type\":\"PropertyValueSpecification\",\"valueRequired\":true,\"valueName\":\"search_term_string\"}}],\"inLanguage\":\"en-US\"},{\"@type\":\"Person\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e\",\"name\":\"Kathryn Royster\",\"image\":{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/image\/\",\"url\":\"https:\/\/secure.gravatar.com\/avatar\/ac60c28082a9a6538663bfb0e8a3752867648c099100b2aa9409599b67af4b16?s=96&d=mm&r=g\",\"contentUrl\":\"https:\/\/secure.gravatar.com\/avatar\/ac60c28082a9a6538663bfb0e8a3752867648c099100b2aa9409599b67af4b16?s=96&d=mm&r=g\",\"caption\":\"Kathryn Royster\"},\"url\":\"https:\/\/dornsife.usc.edu\/wrigley\/author\/kroyster\/\"}]}<\/script>\n<!-- \/ Yoast SEO plugin. -->","yoast_head_json":{"title":"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies","description":"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.","robots":{"index":"index","follow":"follow","max-snippet":"max-snippet:-1","max-image-preview":"max-image-preview:large","max-video-preview":"max-video-preview:-1"},"canonical":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/","og_locale":"en_US","og_type":"article","og_title":"Karen Lloyd, who studies deep ocean microbes, joins USC faculty","og_description":"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.","og_url":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/","og_site_name":"Wrigley Institute for Environment and Sustainability","article_published_time":"2025-02-27T21:52:39+00:00","article_modified_time":"2026-04-02T22:51:53+00:00","og_image":[{"width":2560,"height":1920,"url":"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg","type":"image\/jpeg"}],"author":"Kathryn Royster","twitter_card":"summary_large_image","schema":{"@context":"https:\/\/schema.org","@graph":[{"@type":"Article","@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#article","isPartOf":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/"},"author":{"name":"Kathryn Royster","@id":"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e"},"headline":"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies","datePublished":"2025-02-27T21:52:39+00:00","dateModified":"2026-04-02T22:51:53+00:00","mainEntityOfPage":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/"},"wordCount":12,"commentCount":0,"image":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage"},"thumbnailUrl":"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg","keywords":["climate and carbon management initiative","Earth sciences","faculty","microbiology","oceanography"],"articleSection":["Research"],"inLanguage":"en-US","potentialAction":[{"@type":"CommentAction","name":"Comment","target":["https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#respond"]}]},{"@type":"WebPage","@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/","url":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/","name":"Karen Lloyd, who studies deep ocean microbes, joins USC faculty","isPartOf":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/#website"},"primaryImageOfPage":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage"},"image":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage"},"thumbnailUrl":"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg","datePublished":"2025-02-27T21:52:39+00:00","dateModified":"2026-04-02T22:51:53+00:00","author":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e"},"description":"Biogeochemist Karen Lloyd, who studies deep ocean microbes, has joined the USC faculty as the Wrigley Chair in Environmental Studies.","breadcrumb":{"@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#breadcrumb"},"inLanguage":"en-US","potentialAction":[{"@type":"ReadAction","target":["https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/"]}]},{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#primaryimage","url":"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg","contentUrl":"https:\/\/dornsife.usc.edu\/wrigley\/wp-content\/uploads\/sites\/116\/2025\/02\/IMG_2159-scaled.jpeg","width":2560,"height":1920,"caption":"Karen Lloyd, wearing a dark blue beanie that says \"USC Wrigley Institute,\" smiles as she stands in a craggy, rocky landscape"},{"@type":"BreadcrumbList","@id":"https:\/\/dornsife.usc.edu\/wrigley\/2025\/02\/27\/biogeochemist-karen-lloyd-studies-deep-ocean-microbes-joins-wrigley-institute-usc-faculty\/#breadcrumb","itemListElement":[{"@type":"ListItem","position":1,"name":"Home","item":"https:\/\/dornsife.usc.edu\/wrigley\/"},{"@type":"ListItem","position":2,"name":"Biogeochemist Karen Lloyd joins USC faculty as Wrigley Chair in Environmental Studies"}]},{"@type":"WebSite","@id":"https:\/\/dornsife.usc.edu\/wrigley\/#website","url":"https:\/\/dornsife.usc.edu\/wrigley\/","name":"Wrigley Institute for Environment and Sustainability","description":"Creating a more sustainable and environmentally just future for our planet and all who live on it","potentialAction":[{"@type":"SearchAction","target":{"@type":"EntryPoint","urlTemplate":"https:\/\/dornsife.usc.edu\/wrigley\/?s={search_term_string}"},"query-input":{"@type":"PropertyValueSpecification","valueRequired":true,"valueName":"search_term_string"}}],"inLanguage":"en-US"},{"@type":"Person","@id":"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/8d145e5d8f50e29acee6ca512632733e","name":"Kathryn Royster","image":{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/dornsife.usc.edu\/wrigley\/#\/schema\/person\/image\/","url":"https:\/\/secure.gravatar.com\/avatar\/ac60c28082a9a6538663bfb0e8a3752867648c099100b2aa9409599b67af4b16?s=96&d=mm&r=g","contentUrl":"https:\/\/secure.gravatar.com\/avatar\/ac60c28082a9a6538663bfb0e8a3752867648c099100b2aa9409599b67af4b16?s=96&d=mm&r=g","caption":"Kathryn Royster"},"url":"https:\/\/dornsife.usc.edu\/wrigley\/author\/kroyster\/"}]}},"_links":{"self":[{"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/posts\/3687","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/users\/182"}],"replies":[{"embeddable":true,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/comments?post=3687"}],"version-history":[{"count":12,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/posts\/3687\/revisions"}],"predecessor-version":[{"id":6109,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/posts\/3687\/revisions\/6109"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/media\/3688"}],"wp:attachment":[{"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/media?parent=3687"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/categories?post=3687"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/dornsife.usc.edu\/wrigley\/wp-json\/wp\/v2\/tags?post=3687"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}