{"id":2137,"date":"2025-07-10T23:04:31","date_gmt":"2025-07-10T23:04:31","guid":{"rendered":"https:\/\/dornsife.usc.edu\/heb\/?page_id=2137"},"modified":"2025-08-28T19:50:19","modified_gmt":"2025-08-28T19:50:19","slug":"graduate-program","status":"publish","type":"page","link":"https:\/\/dornsife.usc.edu\/heb\/graduate-program\/","title":{"rendered":"Graduate Program"},"content":{"rendered":"\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--featured-block \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--featured-block\"\n    \n      >\n\n    \n  <div class=\"text-image-container\">\n    <div class=\"text-container\">\n\n                  \n<div class=\"f--field f--eyebrow\">\n\n    \n  <span>PhD Program<\/span>\n\n\n<\/div>\n      \n                  \n<div class=\"f--field f--section-title\">\n\n    \n  <h2>\n          Integrative and Evolutionary Biology Graduate Program\n      <\/h2>\n\n\n<\/div>\n      \n                  \n<div class=\"f--field f--description\">\n\n    \n  <p>The Integrative and Evolutionary Biology (IEB) PhD program of study provides each student with a broad fundamental background in integrative and evolutionary biology with in-depth specialization in one (or more) of research areas. Broad research areas include, but are not limited to evolutionary biology, metabolism and nutrition, neuroscience, endocrinology, glycemic control, energy balance, biomechanics, biological anthropology, skeletal muscle physiology, kinesiology, and conservation.<\/p>\n<p>The curriculum and learning experiences for each student are tailored to their line of research.\u00a0Students take a written and oral screening exam in the first year and a written and oral qualifying exam typically by the end of the fifth semester of study.<\/p>\n<p>For more information on the degree requirements associated with the IEB PhD program, please visit the\u00a0<a href=\"https:\/\/catalogue.usc.edu\/preview_program.php?catoid=21&amp;poid=29641&amp;returnto=8873\">USC catalog<\/a>.<\/p>\n<p>This degree is under the jurisdiction of the Graduate School. Refer to the\u00a0<a href=\"https:\/\/catalogue.usc.edu\/content.php?catoid=21&amp;navoid=8598\">Requirements for Graduation<\/a>\u00a0section and\u00a0<a href=\"https:\/\/catalogue.usc.edu\/content.php?catoid=21&amp;navoid=8860\">The Graduate School<\/a>\u00a0section of this catalogue for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.<\/p>\n\n\n\n<\/div>\n      \n                  \n<div class=\"f--field f--link\">\n\n    \n    \n  \n<a \n  class=\"link\"\n  href= https:\/\/gradadm.usc.edu\/domestic-students\/how-to-apply\/\n    aria-label=\"Read more about Apply Now\"  \n>\n    Apply Now \n  <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" width=\"25\" height=\"25\" xml:space=\"preserve\"><polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#000\" points=\"19.3,27.5 29.3,17.5,19.3,7.5 16.3,10.4 21.4,15.4 6.7,15.4 6.7,19.6 21.4,19.6 16.3,24.6 \"\/><\/svg>\n<\/a>\n\n\n<\/div>\n\n          <\/div>\n\n          <div class=\"image-container\">\n            \n<div class=\"f--field f--image\">\n\n    \n    \n    \n    \n    \n    \n              \n      <img\n                            data-src=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/07\/IEB-Feature-768x768.png\"\n          data-srcset=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/07\/IEB-Feature-1280x1280.png 1280w,https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/07\/IEB-Feature-768x768.png 768w\"          data-sizes=\"(min-width:1024px) 50vw, (min-width:768px) 100vw, 100vw\"          class=\"lazyload\"\n        \n                  role=\"none\"\n        \n        \n                                      \/>\n\n    \n    \n  \n  \n\n<\/div>\n      <\/div>\n    \n  <\/div>\n\n\n  <\/div><\/div>\n\n\n\n      \n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--full-width-cta \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--full-width-cta\"\n    \n      >\n\n    \n<div class=\"inner-wrapper\">\n      <div class=\"header-container\">\n  \n                  \n<div class=\"f--field f--section-title\">\n\n    \n  <h2>\n          Admission Requirements\n      <\/h2>\n\n\n<\/div>\n                        \n<div class=\"f--field f--description\">\n\n    \n  <p>Applicants must have a bachelor&#8217;s degree from an accredited four-year college or university, preferably in an area relevant to the research interests in IEB (e.g. biology, bioanthropology, biomedical engineering, psychology, etc.). <em>GRE is not required<\/em>. <\/p>\n<p>Applicants are evaluated by:<\/p>\n<p>1. Transcripts and GPA<br \/>\n2. Three letters of recommendation<br \/>\n3. A statement of purpose<br \/>\n4. Relevant research experience<\/p>\n<p>A faculty member must serve as the sponsor of admission to the IEB program and advisor throughout the student&#8217;s course of study, We encourage applicants to reach out to potential faculty mentors in advance of submitting their application materials.<\/p>\n\n\n\n<\/div>\n          <\/div>\n    \n      <div class=\"links-container\">\n        \n            \n<div class=\"f--field f--link\">\n\n    \n    \n  \n<a \n  class=\"link\"\n  href= https:\/\/gradadm.usc.edu\/domestic-students\/how-to-apply\/\n    aria-label=\"Read more about Apply Now\"  \n>\n    Apply Now \n  <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" width=\"25\" height=\"25\" xml:space=\"preserve\"><polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#FC0\" points=\"19.3,27.5 29.3,17.5,19.3,7.5 16.3,10.4 21.4,15.4 6.7,15.4 6.7,19.6 21.4,19.6 16.3,24.6 \"\/><\/svg>\n<\/a>\n\n\n<\/div>\n        \n            \n<div class=\"f--field f--link\">\n\n    \n    \n  \n<a \n  class=\"link\"\n  href= https:\/\/dornsife.usc.edu\/heb\/tenure-faculty\/\n    aria-label=\"Read more about Research Faculty\"  \n>\n    Research Faculty \n  <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" width=\"25\" height=\"25\" xml:space=\"preserve\"><polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#FC0\" points=\"19.3,27.5 29.3,17.5,19.3,7.5 16.3,10.4 21.4,15.4 6.7,15.4 6.7,19.6 21.4,19.6 16.3,24.6 \"\/><\/svg>\n<\/a>\n\n\n<\/div>\n        \n            \n<div class=\"f--field f--link\">\n\n    \n    \n  \n<a \n  class=\"link\"\n  href= https:\/\/dornsife.usc.edu\/heb\/heb-faculty-research\/\n    aria-label=\"Read more about Research Labs\"  \n>\n    Research Labs \n  <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" width=\"25\" height=\"25\" xml:space=\"preserve\"><polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#FC0\" points=\"19.3,27.5 29.3,17.5,19.3,7.5 16.3,10.4 21.4,15.4 6.7,15.4 6.7,19.6 21.4,19.6 16.3,24.6 \"\/><\/svg>\n<\/a>\n\n\n<\/div>\n          <\/div>\n  <\/div>\n\n\n\n  <\/div><\/div>\n\n\n\n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n      \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <h2><strong>Financial Support<\/strong><\/h2>\n<p>The program supports each student we admit in the form of teaching assistantships, research assistantships, and fellowships. All awards include full tuition remission, a monthly stipend for living expenses, payment of student health\/dental insurance and other University-wide benefits.<\/p>\n<p>There may also be University requirements for international students. To view these requirements, please visit this\u00a0<a href=\"https:\/\/gradadm.usc.edu\/lightboxes\/international-students-country-requirements\/\">website<\/a>.<\/p>\n<p>USC expects students to communicate effectively in English by demonstrating proficiency in listening, speaking, reading and writing English. <a href=\"http:\/\/ali.usc.edu\/requirement\/\">Click here to review these requirements<\/a>.<\/p>\n<p>For any questions, please contact the Graduate Programs Manager, Doug Burleson (<strong>burleson@usc.edu<\/strong>) or the IEB Student Services Advisor, Adolfo Dela Rosa (<strong>adolfode@usc.edu<\/strong>).<\/p>\n\n\n\n<\/div>\n\n\n  <\/div><\/div>\n\n\n\n\n\n  \n  \n  \t\t              \t\n  \n                                              \n          \n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--news-by-topics \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--news-by-topics\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n\n    <div class=\"heading-container\">\n                  \n<div class=\"f--field f--section-title\">\n\n    \n  <h2>\n          See what our students are up to!\n      <\/h2>\n\n\n<\/div>\n      \n      <div class=\"pagination-wrapper\">\n        <ul class=\"news-by-topics-pagination\"><\/ul>\n      <\/div>\n\n      <div class=\"select-container\">\n        <label class=\"visually-hidden\" for=\"swiper-696560770\">Select option to display slide<\/label>\n        <select name=\"swiper-696560770\">\n                      <option value=\"0\">Graduate Studies<\/option>\n                  <\/select>\n        <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" xml:space=\"preserve\">\n          <polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#FC0\" points=\"7.5,14.9 17.5,24.9 27.5,14.9 24.6,12 17.5,19.2 10.4,12 \"\/>\n        <\/svg>\n      <\/div>\n    <\/div>\n\n    <div class=\"slides-container\">\n              <div class=\"swiper\">\n          <div class=\"swiper-wrapper\">\n                          <div class=\"swiper-slide\">\n\n                <h3 class=\"pane-title\">\n                  Graduate Studies\n                <\/h3>\n\n                                  <div class=\"cards-container\">\n                                          <div class=\"card\">\n                                                  <a href=\"https:\/\/dornsife.usc.edu\/heb\/2025\/09\/02\/marisol-fermin-flores-awarded-the-chancellors-doctoral-incentive-program-cdip-fellowship\/\" target=\"\">\n                                                                                  \n<div class=\"f--field f--image\">\n\n    \n    \n    \n    \n    \n    \n              \n      <img\n                            data-src=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/09\/Marisol-Fermin-Flores-Award-768x432.png\"\n          data-srcset=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/09\/Marisol-Fermin-Flores-Award-1280x720.png 1280w,https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/09\/Marisol-Fermin-Flores-Award-768x432.png 768w\"          data-sizes=\"(min-width:1024px) 23vw, (min-width:768px) 39vw, 83vw\"          class=\"lazyload\"\n        \n                  role=\"none\"\n        \n        \n                                      \/>\n\n    \n    \n  \n  \n\n<\/div>\n                          \n                                                          \n<div class=\"f--field f--cta-title\">\n\n    \n  <h3>\n          Marisol Fermin Flores awarded the Chancellor&#8217;s Doctoral Incentive Program (CDIP) Fellowship\n      <\/h3>\n\n\n<\/div>\n                                                                            <\/a>\n                        \n                      <\/div>\n                                          <div class=\"card\">\n                                                  <a href=\"https:\/\/dornsife.usc.edu\/heb\/2025\/08\/28\/alexander-bashaw-receives-predoctoral-individual-national-research-service-award-from-nih\/\" target=\"\">\n                                                                                  \n<div class=\"f--field f--image\">\n\n    \n    \n    \n    \n    \n    \n              \n      <img\n                            data-src=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Zander-Bashaw-NRSA-Award-768x432.png\"\n          data-srcset=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Zander-Bashaw-NRSA-Award-1280x720.png 1280w,https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Zander-Bashaw-NRSA-Award-768x432.png 768w\"          data-sizes=\"(min-width:1024px) 23vw, (min-width:768px) 39vw, 83vw\"          class=\"lazyload\"\n        \n                  role=\"none\"\n        \n        \n                                      \/>\n\n    \n    \n  \n  \n\n<\/div>\n                          \n                                                          \n<div class=\"f--field f--cta-title\">\n\n    \n  <h3>\n          Alexander Bashaw receives Predoctoral Individual National Research Service Award from NIH\n      <\/h3>\n\n\n<\/div>\n                                                                            <\/a>\n                        \n                      <\/div>\n                                          <div class=\"card\">\n                                                  <a href=\"https:\/\/dornsife.usc.edu\/heb\/2025\/08\/28\/logan-tierno-lauer-receives-predoctoral-individual-national-research-service-award-from-nih\/\" target=\"\">\n                                                                                  \n<div class=\"f--field f--image\">\n\n    \n    \n    \n    \n    \n    \n              \n      <img\n                            data-src=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Logan-Lauer-NRSA-Award-768x432.png\"\n          data-srcset=\"https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Logan-Lauer-NRSA-Award-1280x720.png 1280w,https:\/\/dornsife.usc.edu\/heb\/wp-content\/uploads\/sites\/77\/2025\/08\/Logan-Lauer-NRSA-Award-768x432.png 768w\"          data-sizes=\"(min-width:1024px) 23vw, (min-width:768px) 39vw, 83vw\"          class=\"lazyload\"\n        \n                  role=\"none\"\n        \n        \n                                      \/>\n\n    \n    \n  \n  \n\n<\/div>\n                          \n                                                          \n<div class=\"f--field f--cta-title\">\n\n    \n  <h3>\n          Logan Tierno Lauer receives Predoctoral Individual National Research Service Award from NIH\n      <\/h3>\n\n\n<\/div>\n                                                                            <\/a>\n                        \n                      <\/div>\n                                      <\/div>\n                \n                                      \n<div class=\"f--field f--link\">\n\n    \n    \n  \n<a \n  class=\"link\"\n  href= https:\/\/dornsife.usc.edu\/heb\/graduate-studies-news\/\n    aria-label=\"Read more about More Graduate Studies News\"  \n>\n    More Graduate Studies News \n  <svg version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" width=\"25\" height=\"25\" xml:space=\"preserve\"><polygon fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#FC0\" points=\"19.3,27.5 29.3,17.5,19.3,7.5 16.3,10.4 21.4,15.4 6.7,15.4 6.7,19.6 21.4,19.6 16.3,24.6 \"\/><\/svg>\n<\/a>\n\n\n<\/div>\n                \n              <\/div>\n                      <\/div>\n        <\/div>\n          <\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n","protected":false},"excerpt":{"rendered":"","protected":false},"author":916,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_acf_changed":false,"footnotes":""},"class_list":["post-2137","page","type-page","status-publish","hentry"],"acf":[],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.1.1 - https:\/\/yoast.com\/product\/yoast-seo-wordpress\/ -->\n<title>Graduate Program - Human and Evolutionary Biology<\/title>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/dornsife.usc.edu\/heb\/graduate-program\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Graduate Program - Human and Evolutionary Biology\" \/>\n<meta property=\"og:url\" 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